In 1959, GE Hutchinson posed one of ecology’s most important and enduring questions: Why are there so many species? Answers to this question have a new urgency because of human-driven environmental change. New questions we address are: how will multiple environmental changes affect species diversity, and what are the consequences of losing species for ecosystems? How do the numbers and types of environmental change factors (nutrients, water, temperature, toxins, etc.) depend on each other, and what minimum level of complexity do we need to understand in order to predict species loss?

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