Prof Stan Harpole

Head of Research Group; iDiv Speaker
Prof. Stan Harpole


iDiv publications


Virtanen, Risto, Harpole, W. Stanley, Dunker, Susanne, Eskelinen, Anu

(2024): Multiple global change factors cause declines of a temperate bryophyte. Plant Ecology & Diversity

Eskelinen, Anu, Jessen, Maria-Theresa, Bahamonde, Hector A., Bakker, Jonathan D., Borer, Elizabeth T., Caldeira, Maria C., Harpole, W. Stanley, Jia, Meiyu, Lannes, Luciola S., Nogueira, Carla, Olde Venterink, Harry, Peri, Pablo L., Porath-Krause, Anita J., Seabloom, Eric W., Schroeder, Katie, Tognetti, Pedro M., Yasui, Simone-Louise E., Virtanen, Risto, Sullivan, Lauren L.

(2023): Herbivory and nutrients shape grassland soil seed banks. Nature Communications

Jessen, M.-T., Auge, H., Harpole, W. S., Eskelinen, A.   

(2023): Litter accumulation, not light limitation, drives early plant recruitment. Journal of Ecology

Chen, Qingqing, Wang, Shaopeng, Borer, Elizabeth T., Bakker, Jonathan D., Seabloom, Eric W., Harpole, W. Stanley, Eisenhauer, Nico, Lekberg, Ylva, Buckley, Yvonne M., Catford, Jane A., Roscher, Christiane, Donohue, Ian, Power, Sally A., Daleo, Pedro, Ebeling, Anne, Knops, Johannes M. H., Martina, Jason P., Eskelinen, Anu, Morgan, John W., Risch, Anita C., Caldeira, Maria C., Bugalho, Miguel N., Virtanen, Risto, Barrio, Isabel C., Niu, Yujie, Jentsch, Anke, Stevens, Carly J., Gruner, Daniel S., MacDougall, Andrew S., Alberti, Juan, Hautier, Yann

(2023): Multidimensional responses of grassland stability to eutrophication. Nature Communications

Hallett, Lauren M., Aoyama, Lina, Barabás, György, Gilbert, Benjamin, Larios, Loralee, Shackelford, Nancy, Werner, Chhaya M., Godoy, Oscar, Ladouceur, Emma R., Lucero, Jacob E., Weiss-Lehman, Christopher P., Chase, Jonathan M., Chu, Chengjin, Harpole, W. Stanley, Mayfield, Margaret M., Faist, Akasha M., Shoemaker, Lauren G.

(2023): Restoration ecology through the lens of coexistence theory. Trends in Ecology & Evolution

Ladouceur, E., Isbell, F., Clark, A. T., Harpole, W. S., Reich, P. B., Tilman, G. D., Chase, J. M.

(2023): The recovery of plant community composition following passive restoration across spatial scales. Journal of Ecology

Castro Sánchez-Bermejo, Pablo, Davrinche, Andréa, Matesanz, Silvia, Harpole, W. Stanley, Haider, Sylvia

(2023): Within-individual leaf trait variation increases with phenotypic integration in a subtropical tree diversity experiment. New Phytologist

Hornick, T., Richter, A., Harpole, W. S., Bastl, M., Bohlmann, S., Bonn, A., Bumberger, J., Dietrich, P., Gemeinholzer, B., Grote, R., Heinold, B., Keller, A., Luttkus, M. L., Mader, P., Svara, E. M., Passonneau, S., Punyasena, S. W., Rakosy, D., Richter, R., Sickel, W., Steffan-Dewenter, I., Theodorou, P., Treudler, R., Werchan, B., Werchan, M., Wolke, R., Dunker, S.

(2022): An integrative environmental pollen diversity assessment and its importance for the Sustainable Development Goals. Plants People Planet

Xenophontos, C., Harpole, W. S., Küsel, K. and Clark, A. T.

(2022): Cheating Promotes Coexistence in a Two-Species One-Substrate Culture Model. Frontiers in Ecology and Evolution

Mühlbauer, L. K., Harpole, W. S., Clark, A. T.

(2022): Differences in initial abundances reveal divergent dynamic structures in Gause's predator–prey experiments. Ecology and Evolution

Jessen, M.-T., Auge, H., Harpole, W. S., Hautier, Y., Eskelinen, A.

(2022): Grazing and light modify Silene latifolia responses to nutrients and future climate. PLOS ONE

Virtanen, R., Bakker, J. D., Jessen, M.-T., Sullivan, L. L., Harpole, W. S. and Eskelinen, A.

(2022): Is the bryophyte soil diaspore bank buffered against nutrient enrichment and grazing exclusion?. Plant and Soil

Ladouceur, Emma, Shackelford, Nancy, Bouazza, Karma, Brudvig, Lars, Bucharova, Anna, Conradi, Timo, Erickson, Todd E., Garbowski, Magda, Garvy, Kelly, Harpole, W. Stanley, Jones, Holly P., Knight, Tiffany, Nsikani, Mlungele M., Paterno, Gustavo, Suding, Katharine, Temperton, Vicky M., Török, Péter, Winkler, Daniel E., Chase, Jonathan M.

(2022): Knowledge sharing for shared success in the decade on ecosystem restoration. Ecological Solutions and Evidence

Eskelinen, A., Harpole, W. S., Jessen, M.-T., Virtanen, R., Hautier, Y.

(2022): Light competition drives herbivore and nutrient effects on plant diversity. Nature

Ladouceur, E., Blowes, S. A., Chase, J. M., Clark, A. T., Garbowski, M., Alberti, J., Arnillas, C. A., Bakker, J. D., Barrio, I. C., Bharath, S., Borer, E. T., Brudvig, L. A., Cadotte, M. W., Chen, Q., Collins, S. L., Dickman, C. R., Donohue, I., Du, G., Ebeling, A., Eisenhauer, N., Fay, P. A., Hagenah, N., Hautier, Y., Jentsch, A., Jónsdóttir, I. S., Komatsu, K., MacDougall, A., Martina, J. P., Moore, J. L., Morgan, J. W., Peri, P. L., Power, Sally A., Ren, Z., Risch, A. C., Roscher, C., Schuchardt, Max A., Seabloom, E. W., Stevens, C. J., Veen, G. F., Virtanen, R., Wardle, G. M., Wilfahrt, P. A., Harpole, W. S.

(2022): Linking changes in species composition and biomass in a globally distributed grassland experiment. Ecology Letters

Gill, A. L., Adler, P. B., Borer, E. T., Buyarski, C. R., Cleland, E. E., D'Antonio, C. M., Davies, K. F., Gruner, D. S., Harpole, W. S., Hofmockel, K. S., MacDougall, A. S., McCulley, R. L., Melbourne, B. A., Moore, J. L., Morgan, J. W., Risch, A. C., Schütz, M., Seabloom, E. W., Wright, J. P., Yang, L. H. and Hobbie, S. E.

(2022): Nitrogen increases early-stage and slows late-stage decomposition across diverse grasslands. Journal of Ecology

Muehleisen, A. J., Watkins, C. R. E., Altmire, G. R., Shaw, E. A., Case, M. F., Aoyama, L., Brambila, A., Reed, P. B., LaForgia, M., Borer, E. T., Seabloom, E. W., Bakker, J. D., Amillas, C. A., Biederman, L., Chen, Q., Cleland, E. E., Fay, P. A., Hagenah, N., Harpole, S., Hautier, Y., Henning, J. A., Knops, J. M. H., Komatsu, K. J., Ladouceur, E., MacDougall, A., McCulley, R. L., Moore, J. L., Ohlert, T., Power, S. A., Stevens, C. J., Wilfahrt, P., Hallett, L. M.

(2022): Nutrient addition drives declines in grassland species richness primarily via enhanced species loss. Journal of Ecology

Dunker, S., Boyd, M., Durka, W., Erler, S., Harpole, W. S., Henning, S., Herzschuh, U., Hornick, T., Knight, T., Lips, S., Mäder, P., Švara, E. M., Mozarowski, S., Rakosy, D., Römermann, C., Schmitt-Jansen, M., Stoof-Leichsenring, K., Stratmann, F., Treudler, R., Virtanen, R., Wendt-Potthoff, K.Wilhelm, C.

(2022): The potential of multispectral imaging flow cytometry for environmental monitoring. Cytometry Part A

Wilfahrt, P. A., Asmus, A. L., Seabloom, E. W., Henning, J. A., Adler, P., Arnillas, C. A., Bakker, J. D., Biederman, L., Brudvig, L. A., Cadotte, M., Daleo, P., Eskelinen, A., Firn, J., Harpole, W. S., Hautier, Y., Kirkman, K. P., Komatsu, K. J., Laungani, R., MacDougall, A., McCulley, R. L., Moore, J. L., Morgan, J. W., Mortensen, B., Ochoa Hueso, R., Ohlert, T., Power, S. A., Price, J., Risch, A. C., Schuetz, M., Shoemaker, L., Stevens, C., Strauss, A. T., Tognetti, P. M., Virtanen, R., Borer, E. T.

(2021): Temporal rarity is a better predictor of local extinction risk than spatial rarity. Ecology

A. Compagnoni, S. Levin, D. Z. Childs, S. Harpole, M. Paniw, G. Römer, J. H. Burns, J. Che-Castaldo, N. Rüger, G. Kunstler, J. M. Bennett, C. R. Archer, O. R. Jones, R. Salguero-Gómez, T. M. Knight

(2021): Herbaceous perennial plants with short generation time have stronger responses to climate anomalies than those with longer generation time. Nature Communications

A. T. Clark, J.-F. Arnoldi, Y. R. Zelnik, G. Barabas, D. Hodapp, C. Karakoç, S. König, V. Radchuk, I. Donohue, A. Huth, C. Jacquet, C. de Mazancourt, A. Mentges, D. Nothaaß, L. G. Shoemaker, F. Taubert, T. Wiegand, S. Wang, J. M. Chase, M. Loreau, S. Harpole

(2021): General statistical scaling laws for stability in ecological systems. Ecology Letters

C. Xenophontos, M. Taubert, W. S. Harpole, K. Küsel

(2021): Phylogenetic and metabolic diversity have contrasting effects on the ecological functioning of bacterial communities. FEMS Microbiology Ecology

E. W. Seabloom, E. Batzer, J. M. Chase, W. Stanley Harpole, P. B. Adler, S. Bagchi, J. D. Bakker, I. C. Barrio, L. Biederman, E. H. Boughton, M. N. Bugalho, M. C. Caldeira, J. A. Catford, P. Daleo, N. Eisenhauer, A. Eskelinen, S. Haider, L. M. Hallett, I. Svala Jónsdóttir, K. Kimmel, M. Kuhlman, A. MacDougall, C. D. Molina, J. L. Moore, J. W. Morgan, R. Muthukrishnan, T. Ohlert, A. C. Risch, C. Roscher, M. Schütz, G. Sonnier, P. M. Tognetti, R. Virtanen, P. A. Wilfahrt, E. T. Borer

(2021): Species loss due to nutrient addition increases with spatial scale in global grasslands. Ecology Letters

H. Hillebrand, I. Donohue, W. S. Harpole, D. Hodapp, M. Kucera, A. M. Lewandowska, J. Merder, J. M. Montoya, J. A. Freund

(2021): Reply to: Empirical pressure-response relations can benefit assessment of safe operating spaces. Nature Ecology & Evolution

L. Korell, H. Auge, J. M. Chase, W. S. Harpole, T. M. Knight

(2021): Responses of plant diversity to precipitation change are strongest at local spatial scales and in drylands. Nature Communications

M. E. Wilcots, W. S. Harpole, E. W. Seabloom, E. T. Borer

(2021): Community change can buffer chronic nitrogen impacts, but multiple nutrients tip the scale. Ecology

P. Hofmann, A. Clark, P. Hoffmann, A. Chatzinotas, W. S. Harpole, S. Dunker

(2021): Beyond nitrogen: phosphorus – estimating the minimum niche dimensionality for resource competition between phytoplankton. Ecology Letters

Seabloom, E. W., P. B. Adler, J. Alberti, L. Biederman, Y. M. Buckley, M. W. Cadotte, S. L. Collins, L. Dee, P. A. Fay, J. Firn, N. Hagenah, W. S. Harpole, Y. Hautier, A. Hector, S. E. Hobbie, F. Isbell, J. M. H. Knops, K. J. Komatsu, R. Laungani, A. MacDougall, R. L. McCulley, J. L. Moore, J. W. Morgan, T. Ohlert, S. M. Prober, A. C. Risch, M. Schuetz, C. J. Stevens, E. T. Borer

(2021): Increasing effects of chronic nutrient enrichment on plant diversity loss and ecosystem productivity over time. Ecology

Borer, E. T., W. S. Harpole, P. B. Adler, C. A. Arnillas, M. N. Bugalho, M. W. Cadotte, M. C. Caldeira, S. Campana, C. R. Dickman, T. L. Dickson, I. Donohue, A. Eskelinen, J. L. Firn, P. Graff, D. S. Gruner, R. W. Heckman, A. M. Koltz, K. J. Komatsu, L. S. Lannes, A. S. MacDougall, J. P. Martina, J. L. Moore, B. Mortensen, R. Ochoa-Hueso, H. Olde Venterink, S. A. Power, J. N. Price, A. C. Risch, M. Sankaran, M. Schütz, J. Sitters, C. J. Stevens, R. Virtanen, P. A. Wilfahrt, E. W. Seabloom

(2020): Nutrients cause grassland biomass to outpace herbivory. Nature Communications

Broadbent, A. A. D., Firn, J., McGree, J. M., Borer, E. T., Buckley, Y. M., Harpole, W. S., ..., Eisenhauer, N. et al

(2020): Dominant native and non-native graminoids differ in key leaf traits irrespective of nutrient availability. Global Ecology and Biogeography

Clark, A. T., Ann Turnbull, L., Tredennick, A., Allan, E., Harpole, W. S., Mayfield, M. M., Soliveres, S., Barry, K., Eisenhauer, N., de Kroon, H., Rosenbaum, B., Wagg, C., Weigelt, A., Feng, Y., Roscher, C., Schmid, B.

(2020): Predicting species abundances in a grassland biodiversity experiment: Trade-offs between model complexity and generality. Journal of Ecology

Eskelinen, A., K. Gravuer, S. Harpole, S. Harrison, R. Virtanen and Y. Hautier

(2020): Resource-enhancing global changes drive a whole-ecosystem shift to faster cycling but decrease diversity. Ecology

Gilbert, B., MacDougall, A. S., Kadoya, T., Akasaka, M., Bennett, J. R., ..., Harpole, W. S., ..., Fay, P. A.

(2020): Climate and local environment structure asynchrony and the stability of primary production in grasslands. Global Ecology and Biogeography

Hillebrand, H., Donohue, I., Harpole, W. S., Hodapp, D., Kucera, M., Lewandowska, A. M., Merder, J., Montoya, J. M., Freund, J. A.

(2020): Thresholds for ecological responses to global change do not emerge from empirical data. Nature Ecology & Evolution

Korell, L., Auge, H., Chase, J. M., Harpole, W. S., Knight, T. M.

(2020): We need more realistic climate change experiments for understanding ecosystems of the future. Global Change Biology

Ladouceur, E., Stanley Harpole, W., Blowes, S. A., Roscher, C., Auge, H., Seabloom, E. W., Chase, J. M.

(2020): Reducing dispersal limitation via seed addition increases species richness but not above-ground biomass. Ecology Letters

Mühlbauer, L. K., M. Schulze, W. S. Harpole, A. T. Clark

(2020): gauseR: Simple methods for fitting Lotka-Volterra models describing Gause’s “Struggle for Existence”. Ecology and Evolution

Weise, H., Auge, H., Baessler, C., Bärlund, I., Bennett, E. M., Berger, U., Bohn, F., Bonn, A., ..., Dunker, S., ..., Groeneveld, J., ..., Harpole, S., ..., Klotz, S., Kühn, I., ..., Seppelt, R., ..., Wirth, C., Wolf, C., Grimm, V.



(2020): Resilience trinity: safeguarding ecosystem functioning and services across three different time horizons and decision contexts. Oikos

Barry, K. E., H. de Kroon, P. Dietrich, W. Stanley Harpole, A. Roeder, B. Schmid, A. T. Clark, M. M. Mayfield, C. Wagg, C. Roscher

(2019): Linking local species coexistence to ecosystem functioning: A conceptual framework from ecological first principles in grassland ecosystems. In: Nico Eisenhauer, David A. Bohan, Alex J. Dumbrell (Eds.) Mechanisms underlying the relationship between biodiversity and ecosystem function Advances in Ecological Research

Clark, A. T., Barry, K. E., Roscher, C., Buchmann, T., Loreau, M., Harpole, W. S.

(2019): How to Estimate Complementarity and Selection Effects from an Incomplete Sample of Species. Methods in Ecology and Evolution

Clark, A. T., H. Hillebrand, W. S. Harpole

(2019): Scale Both Confounds and Informs Characterization of Species Coexistence in Empirical Systems. The American Naturalist

Cleland, E. E., Lind, E. M., DeCrappeo, N. M., DeLorenze, E., Wilkins, R. A., Adler, P. B., Bakker, J. D., Brown, C. S., Davies, K. F., Esch, E., Firn, J., Gressard, S., Gruner, D. S., Hagenah, N., Harpole, W. S., et al.

(2019): Belowground Biomass Response to Nutrient Enrichment Depends on Light Limitation Across Globally Distributed Grasslands. Ecosystems

Firn, J., McGree, J. M., Harvey, E., Flores-Moreno, H., Schütz, M., Buckley, Y. M., Borer, E. T., Seabloom, E. W., La Pierre, K. J., MacDougall, A. M., Prober, S. M., Stevens, C. J., Sullivan, L. L., Porter, E., Ladouceur, E., Allen, C., Moromizato, K. H., Morgan, J. W., Harpole, W. S., Hautier, Y., Eisenhauer, N., ..., Roscher, C., et al.

(2019): Leaf nutrients, not specific leaf area, are consistent indicators of elevated nutrient inputs. Nature Ecology & Evolution

Hofmann, P., Chatzinotas, A., Harpole, W. S., Dunker, S

(2019): Temperature and stoichiometric dependence of phytoplankton traits. Ecology

Korell, L., Auge, H., Chase, J. M., Harpole, W. S., Knight, T. M.

(2019): Understanding plant communities of the future requires filling knowledge gaps. Global Change Biology

Parker, S. S., Harpole, W. S., Seabloom, E. W.

(2019): Plant species natural abundances are determined by their growth and modification of soil resources in monoculture. Plant and Soil

Shoemaker, L. G., Sullivan, L. L., Donohue, I., Cabral, J. S., ...., Chase, J. M., Chu, C., Harpole, W. S., Huth, A., ..., May, F., ..., Wiegand, T., Yang, Q., Abbott, K. C.

(2019): Integrating the underlying structure of stochasticity into community ecology. Ecology

Anderson, T. M., Griffith, D. M., Grace, J. B., Lind, E. M., Adler, P. B., Biederman, L. A., Blumenthal, D. M., Daleo, P., Firn, J., Hagenah, N., Harpole, W. S., et al.

(2018): Herbivory and eutrophication mediate grassland plant nutrient responses across a global climatic gradient. Ecology

Bannar-Martin, K. H., Kremer, C. T., Ernest, M., Leibold, M. A., Auge, H., Chase, J., Declerck, S. A. J., Eisenhauer, N., Harpole, S., Hillebrand, H., Isbell, F., Koffel, T., Larsen, S., Narwani, A., Petermann, J. S., Roscher, C., Cabral, J. S., Supp, S. R.

(2018): Integrating community assembly and biodiversity to better understand ecosystem function: the Community Assembly and the Functioning of Ecosystems (CAFE) approach. Ecology Letters

Frater, P. N., Borer, E. T., Fay, P. A., Jin, V., Knaeble, B., Seabloom, E., Sullivan, L., Wedin, D. A., Harpole, W. S.

(2018): Nutrients and environment influence arbuscular mycorrhizal colonization both independently and interactively in Schizachyrium scoparium. Plant and Soil

Hillebrand, H., Blasius, B., Borer, E. T., Chase, J. M., Downing, J., Eriksson, B. K., Filstrup, C. T., Harpole, W. S., Hodapp, D., Larsen, S., et al.

(2018): Biodiversity change is uncoupled from species richness trends: consequences for conservation and monitoring. Journal of Applied Ecology

Hodapp, D., Borer, E. T., Harpole, W. S., Lind, E. M., Seabloom, E. W., Adler, P. B., Alberti, J., Arnillas, C. A., Bakker, J. D., Biederman, L., et al.

(2018): Spatial heterogeneity in species composition constrains plant community responses to herbivory and fertilisation. Ecology Letters

Kröhnert, M., Anderson, R., Bumberger, J., Dietrich, P., Harpole, W., Maas, H. G.


Mortensen, B., Danielson, B., Harpole, S. W., Alberti, J., Arnillas, C. A., Biederman, L., Borer, E. T., Cadotte, M. W., Dwyer, J. M., Hagenah, N., Hautier, Y., Peri, P. L., Seabloom, E. W.

(2018): Herbivores safeguard plant diversity by reducing variability in dominance. Journal of Ecology

Borer, E. T., Grace, J. B., Harpole, W. S., MacDougall, A. S., Seabloom, E. W.

(2017): A decade of insights into grassland ecosystem responses to global environmental change. Nature Ecology & Evolution

Ceulemans, T., Bodé, S., Bollyn, J., Harpole, S., Coorevits, K., Peeters, G., Van Acker, K., Smolders, E., Boeckx, P., Honnay, O.

(2017): Phosphorus resource partitioning shapes phosphorus acquisition and plant species abundance in grasslands. Nature Plants

Harpole, W. S., Sullivan, L. L., Lind, E. M., Firn, J., Adler, P. B., Borer, E. T., Chase, J., Fay, P. A., Hautier, Y., Hillebrand, H., et al.

(2017): Out of the shadows: multiple nutrient limitations drive relationships among biomass, light and plant diversity. Functional Ecology

Dunker, S., Nadrowski, K., Jakob, T., Kasprzak, P., Becker, A., Langner, U., Kunath, C., Harpole, S., Wilhelm, C.

(2016): Assessing in situ dominance pattern of phytoplankton classes by dominance analysis as a proxy for realized niches. Harmful Algae

Flores-Moreno, H., Reich, P. B., Lind, E. M., Sullivan, L. L., Seabloom, E. W., Yahdjian, L., MacDougall, A. S., Reichmann, L. G., Alberti, J., Baez, S., Bakker, J. D., Cadotte, M. W., Caldeira, M. C., Chaneton, E. J., D'Antonio, C. M., Fay, P. A., Firn, J., Hagenah, N., Harpole, W. S., et al.

(2016): Climate modifies response of non-native and native species richness to nutrient enrichment. Philosophical Transactions of the Royal Society B - Biological Sciences

Grace, J. B., Anderson, T. M., Seabloom, E. W., Borer, E. T., Adler, P. B., Harpole, W. S., Hautier, Y., Hillebrand, H., Lind, E. M., Pärtel, M., et al.

(2016): Integrative modelling reveals mechanisms linking productivity and plant species richness. Nature

Harpole, W. S.

(2016): The hydrological niche and spatially structured species co-existence. Journal of Vegetation Science

Harpole, W. S., Sullivan, L. L., Lind, E. M., Firn, J., Adler, P. B., Borer, E. T., Chase, J., Fay, P. A., Hautier, Y., Hillebrand, H., et al.

(2016): Addition of multiple limiting resources reduces grassland diversity. Nature

Lewandowska, A. M., Biermann, A., Borer, E. T., Cebrián-Piqueras, M. A., Declerck, S. A. J., Meester, L. D., Donk, E. V., Gamfeldt, L., Gruner, D. S., Hagenah, N., Harpole, W. S., Kirkman, K. P., Klausmeier, C. A., Kleyer, M., Knops, J. M. H., Lemmens, P., Lind, E. M., Litchman, E., Mantilla-Contreras, J., Martens, K., Meier, S., Minden, V., Moore, J. L., Venterink, H. O., Seabloom, E. W., Sommer, U., Striebel, M., Trenkamp, A., Trinogga, J., Urabe, J., Vyverman, W., Waal, D. B. V. d., Widdicombe, C. E., Hillebrand, H.

(2016): The influence of balanced and imbalanced resource supply on biodiversity– functioning relationship across ecosystems. Philosophical Transactions of the Royal Society B

Schuman, M. C., van Dam, N. M., Beran, F., Harpole, W. S.

(2016): How does plant chemical diversity contribute to biodiversity at higher trophic levels?. Current Opinion in Insect Science

Stein, C., Harpole, W. S., Suding, K. N.

(2016): Transitions and invasion along a grazing gradient in experimental California grasslands. Ecology

Sullivan, L. L., Danielson, B. J., Harpole, W. S.

(2016): Mammalian Herbivores Alter the Population Growth and Spatial Establishment of an Early-Establishing Grassland Species. PLoS One

Tredennick, A. T., Adler, P. B., Grace, J. B., Harpole, W. S., Borer, E. T., Seabloom, E. W., Anderson, T. M., Bakker, J. D., Biederman, L. A., Brown, C. S., et al.

(2016): Comment on “Worldwide evidence of a unimodal relationship between productivity and plant species richness”. Science

Fay, P. A., Prober, S. M., Harpole, W. S., Knops, J. M. H., Bakker, J. D., Borer, E. T., Lind, E. M., MacDougall, A. S., Seabloom, E. W., Wragg, P. D., et al.

(2015): Grassland productivity limited by multiple nutrients. Nature Plants

Leff, J. W., Jones, S. E., Prober, S. M., Barberán, A., Borer, E. T., Firn, J. L., Harpole, W. S., Hobbie, S. E., Hofmockel, K. S., Knops, J. M. H., et al.

(2015): Consistent responses of soil microbial communities to elevated nutrient inputs in grasslands across the globe. Proceedings of the National Academy of Sciences

Prober, S. M., Leff, J. W., Bates, S. T., Borer, E. T., Firn, J., Harpole, W. S., Lind, E. M., Seabloom, E. W., Adler, P. B., Bakker, J. D., et al.

(2015): Plant diversity predicts beta but not alpha diversity of soil microbes across grasslands worldwide. Ecology Letters

Seabloom, E. W., Borer, E. T., Buckley, Y. M., Cleland, E. E., Davies, K. F., Firn, J., Harpole, W. S., Hautier, Y., Lind, E. M., MacDougall, A. S., et al.

(2015): Plant species' origin predicts dominance and response to nutrient enrichment and herbivores in global grasslands. Nature Communications

Stevens, C. J., Lind, E. M., Hautier, Y., Harpole, W. S., Borer, E. T., Hobbie, S., Seabloom, E. W., Ladwig, L., Bakker, J. D., Chu, C., et al.

(2015): Anthropogenic nitrogen deposition predicts local grassland primary production worldwide. Ecology
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